
Lab Group
Luke Miller, Ph.D., Principal Investigator

I label myself as a marine ecophysiologist, biomechanic, intertidal ecologist, or just gadget guy, all depending on the day and my mood. I am primarily interested in how marine organisms deal with the physical environment including the weather and waves, as well as how organisms deal with each other via competition and predation. Much of my research has focused on northeastern Pacific rocky intertidal habitats from California to Alaska, but I have also worked on New England rocky shore communities, on Antarctic scallops, and in kelp forests and estuary habitats. I am interested in how individual organisms and communities might cope with the effects of climate change, but also how they cope with day-to-day variation in their environment. I employ a variety of methods and approaches that draw from the fields of biomechanics and physiology, and do work both in the field and lab.
Ph.D. students
M.Sc. students

Bailey McKernan
Bailey graduated from UCSB with a B.S. in Environmental Studies in 2023. She is interested in how organisms survive in dynamic environments and how species interactions may shift under climate change.
Undergraduate students

Miles Ghannadian

Neenah Mendez

Jasmine Pooni

Zainab Al-Jasim
- Andrew Chau
Alumni

Monica Klopp, M.Sc.
Monica completed her Masters degree at San Diego State University in Fall of 2025, where she studied the impacts of water quality changes in Southern California estuaries on the behavior and physiology of Mytilus galloprovincialis mussels.
Klopp, M.J. (2025) Cardiac and valve-gaping activity of mussels in response to common stressors in San Diego estuaries. Masters thesis, San Diego State University. (PDF)
Lily McIntire, Ph.D.
Lily McIntire completed her Ph.D. as part of the Joint Doctoral Program in Ecology with UC Davis in 2025. She joined the Miller Lab after she finished her master’s degree at Cal Poly Humboldt. She is interested in the thermal ecology of intertidal organisms, particularly how elevated temperatures like those caused by climate change will affect these animals.
McIntire, L.C. (2025). Beat the Heat: Examining Behavioral Trade-Offs in the Face of Climate Warming for Intertidal Ectotherms. Ph.D. Dissertation, San Diego State University and U.C. Davis Joint Doctoral Program in Ecology. (PDF)


Lauren Strope, M.Sc.
Lauren Strope earned her Masters degree at San Diego State University in 2023. She studied the growth and survival of the native oyster Ostrea lurida and interactions with eelgress in a restored oyster bed and eelgrass system in Newport Bay, southern California.
Strope, L.Y. (2023) Effects of eelgrass presence and estuarine abiotic factors on oyster physiology. Masters thesis, San Diego State University (PDF)
Gabriella Kalbach
Gabriella graduated from CSU Monterey Bay, and while in the Miller Lab worked on using biosensors to track gaping behavior and heart rates of Crassostrea [Magallana] gigas oysters in southern San Diego Bay and nearby estuaries.

Past Undergraduate Research Assistants
Grace Addleman
Karla Alcaraz
Angel Sanchez
Malea Arriola
Caileen Baldwin
Ismail Siham
Ka Mwee Paw
Aya Kareem
Maia Little
Erin McClure
Aranza Maldonado
Nature Currey
Sebastian Penalosa
Richard Villareal
Trevor Tompkins
Kristen Waterford
McKenna Purdy
Rio Case
Paula Crespo
Arianna Dial
Simone Parsons
Elisabeth Rotschedl
Alani Black
Dorian Marquez